I seem to remember that the reason Muriah's broo gang are said to have resented her is her sterility. While I doubt if this was the whole story, it does sound as if it had *some* effect on them.
Personally I go for the idea that broo can impregnate *anything*. It's just that some things are more difficult than others. Fertile female mammals are easy. Anything progressively harder (infertile female, male, non-mammal, inanimate object, True Dragon) requires a ritual of progressively greater difficulty. This ritual doesn't actually have to involve acute discomfort for the victim in order to work, but broo being broo, it usually does anyway.
My guess would be that feral broo know no such rituals. Broo gangs such as in the Rubble know at least the ones for male humans. And good old Ralzakark knows the lot, of course. The only question is whether he can be bothered.
Jane Williams jane_at_williams.nildram.co.ukhttp://homepages.nildram.co.uk/~janewill/gloranth/index.shtml
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