I'd let people do it if they justified it in their background (or ideally in their 100 words). We know that occasional Orlanthi are born able to fly so that's easy (for Orlanthi - but I'd be less lenient for others).
I think I'd consider requiring:
i) some justification in terms of culture, ii) something in the character background, and iii) appropriate flaws be taken
For example, a character who takes "spit fire" might get "dragonfriend" or "suspected of chaos magic" as a flaw depending on the justification they proposed. [The latter is probably reserved for no justification or a weak one]
e.g., "Verlak gained the spit fire feat from Orlanth Dragonfriend" versus "Verlak can spit fire"
I wouldn't increase the expense.
Thom
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